/* ========================================================================
   CATTLEYAS · CSS de la presentación V1
   Extiende theme.css base, adapta Reveal.js al tema oscuro botánico
   ======================================================================== */

/* === REVEAL.JS THEME OVERRIDE === */

.reveal-viewport {
  background: var(--bg-base) !important;
}

.reveal {
  font-family: var(--font-body);
  font-size: 24px;  /* base scale Reveal */
  font-weight: 400;
  color: var(--ink-secondary);
  -webkit-font-smoothing: antialiased;
}

.reveal .slides {
  text-align: left;
}

.reveal .slides > section,
.reveal .slides > section > section {
  padding: 0;
  font-size: 1em;
  line-height: var(--lh-normal);
}

/* Scrollbar discreto en componentes que pueden hacer scroll interno */
.species-content,
.hybrids-grid,
.compare-wrap,
.history-timeline {
  scrollbar-width: thin;
  scrollbar-color: var(--gold-soft) transparent;
}

.species-content::-webkit-scrollbar,
.hybrids-grid::-webkit-scrollbar { width: 8px; }
.species-content::-webkit-scrollbar-track,
.hybrids-grid::-webkit-scrollbar-track { background: transparent; }
.species-content::-webkit-scrollbar-thumb,
.hybrids-grid::-webkit-scrollbar-thumb {
  background: var(--gold-soft);
  border-radius: 4px;
}
.species-content::-webkit-scrollbar-thumb:hover,
.hybrids-grid::-webkit-scrollbar-thumb:hover {
  background: var(--gold);
}

/* === Tipografía Reveal === */

.reveal h1,
.reveal h2,
.reveal h3,
.reveal h4,
.reveal h5,
.reveal h6 {
  font-family: var(--font-display);
  color: var(--ink-primary);
  font-weight: 700;
  line-height: var(--lh-tight);
  letter-spacing: var(--tracking-tight);
  margin: 0 0 var(--space-md);
  text-transform: none;
}

.reveal h1 { font-size: clamp(3rem, 7vw, 5rem); }
.reveal h2 { font-size: clamp(2.25rem, 5vw, 3.5rem); }
.reveal h3 { font-size: clamp(1.6rem, 3vw, 2.2rem); }
.reveal h4 { font-size: 1.4rem; font-weight: 600; }

.reveal p { margin: 0 0 var(--space-md); }
.reveal a { color: var(--gold-bright); text-decoration: none; }
.reveal a:hover { color: var(--gold); }
.reveal em { color: var(--neon-bright); }

.reveal strong { color: var(--ink-primary); font-weight: 600; }

/* Itálica científica */
.sci, .reveal .sci {
  font-family: var(--font-serif);
  font-style: italic;
  font-weight: 500;
  letter-spacing: 0.005em;
}

/* === CONTROLES DE REVEAL === */

.reveal .controls {
  color: var(--gold);
}
.reveal .controls .navigate-left.enabled,
.reveal .controls .navigate-right.enabled,
.reveal .controls .navigate-up.enabled,
.reveal .controls .navigate-down.enabled {
  opacity: 0.55;
}
.reveal .controls button:hover {
  opacity: 1 !important;
}

.reveal .progress {
  color: var(--gold);
  height: 3px;
  background: rgba(255,255,255,0.06);
}
.reveal .slide-number {
  color: var(--ink-muted);
  background: var(--bg-overlay);
  padding: 4px 10px;
  border-radius: var(--radius-pill);
  font-family: var(--font-body);
  font-size: 12px;
  font-weight: 500;
  letter-spacing: 0.04em;
}

/* === LAYOUT DE SLIDE === */

.slide-pad {
  padding: var(--space-xl) var(--space-2xl);
  max-width: 1380px;
  margin: 0 auto;
  width: 100%;
  box-sizing: border-box;
}

.slide-narrow { max-width: 1080px; }
.slide-full   { max-width: none; padding: var(--space-lg) var(--space-xl); }

/* === PORTADA (slide 1) === */

.cover {
  display: grid;
  grid-template-rows: auto 1fr auto;
  min-height: 100%;
  height: 100%;
  gap: var(--space-md);
  text-align: center;
  padding: var(--space-xl);
  box-sizing: border-box;
  width: 100%;
}

.cover-society {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: var(--space-xs);
}

.cover-hero-logo {
  width: clamp(160px, 22vh, 230px);
  aspect-ratio: 1;
  border-radius: 50%;
  object-fit: cover;
  background: var(--bg-surface);
  margin: 0 auto;
  box-shadow:
    0 0 64px rgba(196,110,224,0.20),
    0 0 0 1px var(--border-default);
  animation: glow-breathe 5s ease-in-out infinite;
}

.cover-center {
  display: flex;
  flex-direction: column;
  align-items: center;
  justify-content: center;
  gap: var(--space-md);
}

.cover-title {
  font-family: var(--font-display);
  font-weight: 700;
  font-size: clamp(2.4rem, 6vw, 4.8rem);
  line-height: 0.95;
  letter-spacing: -0.025em;
  color: var(--ink-primary);
}

.cover-title .roman {
  display: block;
  font-family: var(--font-serif);
  font-style: italic;
  font-weight: 500;
  font-size: 0.55em;
  color: var(--neon-bright);
  margin-top: var(--space-xs);
  letter-spacing: 0;
}

.cover-subtitle {
  font-family: var(--font-serif);
  font-style: italic;
  font-size: clamp(1.15rem, 2vw, 1.6rem);
  color: var(--ink-secondary);
}

.cover-year {
  display: flex;
  gap: var(--space-md);
  align-items: center;
  color: var(--ink-tertiary);
  font-size: 0.85rem;
  letter-spacing: var(--tracking-wide);
}
.cover-year .anchor { color: var(--gold); font-weight: 600; }

.cover-footer {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: var(--space-md);
}

.cover-footer-logos {
  display: flex;
  gap: clamp(var(--space-lg), 4vw, var(--space-2xl));
  align-items: center;
  opacity: 0.7;
  filter: brightness(1.1);
}
.cover-footer-logos img {
  height: clamp(36px, 5vh, 52px);
  background: rgba(245,242,237,0.92);
  padding: 6px 10px;
  border-radius: var(--radius-sm);
}

/* === SECCIÓN SEPARADORES (intro a sección) === */

.section-divider {
  display: grid;
  place-items: center;
  min-height: 100%;
  height: 100%;
  width: 100%;
  text-align: center;
  position: relative;
  overflow: hidden;
  box-sizing: border-box;
}

.section-divider .accent-bg {
  position: absolute;
  inset: 0;
  background:
    radial-gradient(ellipse 40% 50% at 50% 50%, var(--accent-color, var(--gold-soft)), transparent 70%);
  opacity: 0.25;
  z-index: 0;
}

.section-divider-content {
  position: relative;
  z-index: 1;
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: var(--space-md);
  max-width: 800px;
  padding: 0 var(--space-xl);
}

.section-divider-eyebrow {
  font-size: 0.7rem;
  font-weight: 600;
  letter-spacing: var(--tracking-widest);
  text-transform: uppercase;
  color: var(--accent-color, var(--gold));
}

.section-divider-title {
  font-family: var(--font-display);
  font-weight: 700;
  font-size: clamp(2.5rem, 6vw, 5rem);
  line-height: 0.95;
  letter-spacing: -0.02em;
  color: var(--ink-primary);
}

.section-divider-meta {
  font-family: var(--font-serif);
  font-style: italic;
  color: var(--ink-secondary);
  font-size: 1.1rem;
}

/* === ÁRBOL TAXONÓMICO INTERACTIVO === */

.tax-tree {
  font-family: var(--font-body);
  font-size: 0.95rem;
  width: 100%;
  max-width: 1180px;
  margin: 0 auto;
}

.tax-tree-toolbar {
  display: flex;
  gap: var(--space-sm);
  justify-content: flex-end;
  margin-bottom: var(--space-md);
}

.tax-tree-toolbar button {
  background: transparent;
  color: var(--ink-tertiary);
  border: 1px solid var(--border-default);
  padding: 6px 14px;
  font-size: 0.8rem;
  border-radius: var(--radius-pill);
  cursor: pointer;
  font-family: inherit;
  letter-spacing: var(--tracking-wide);
  transition: all var(--t-fast);
}
.tax-tree-toolbar button:hover {
  border-color: var(--border-accent);
  color: var(--ink-primary);
  background: var(--gold-soft);
}

.tax-node {
  margin: var(--space-2xs) 0;
}

.tax-node-header {
  display: grid;
  grid-template-columns: 24px minmax(140px, 220px) minmax(120px, 200px) auto 60px;
  align-items: center;
  gap: var(--space-md);
  padding: var(--space-sm) var(--space-md);
  background: var(--bg-surface);
  border: 1px solid var(--border-subtle);
  border-radius: var(--radius-md);
  cursor: pointer;
  transition: all var(--t-fast);
  border-left-width: 3px;
  border-left-color: var(--node-color, var(--gold));
}
.tax-node-header:hover {
  background: var(--bg-elevated);
  border-color: var(--border-accent);
}

.tax-node-toggle {
  width: 20px;
  height: 20px;
  display: grid;
  place-items: center;
  font-size: 14px;
  color: var(--ink-tertiary);
  transition: transform var(--t-fast);
}

.tax-node[open] > .tax-node-header > .tax-node-toggle {
  transform: rotate(90deg);
  color: var(--gold);
}

.tax-node-toggle::before {
  content: "▶";
}

.tax-node-leaf .tax-node-toggle::before {
  content: "·";
  color: var(--ink-muted);
}

.tax-node-name {
  font-weight: 600;
  color: var(--ink-primary);
}

.tax-node-name.scientific {
  font-family: var(--font-serif);
  font-style: italic;
  font-weight: 500;
}

.tax-node-meta {
  color: var(--ink-tertiary);
  font-size: 0.85em;
}

.tax-node-count {
  text-align: right;
  font-family: var(--font-display);
  font-weight: 700;
  color: var(--node-color, var(--gold));
  font-size: 1.1em;
}

.tax-node-children {
  margin-left: var(--space-xl);
  margin-top: var(--space-xs);
  padding-left: var(--space-md);
  border-left: 1px dashed var(--border-default);
  display: none;
}

.tax-node[open] > .tax-node-children {
  display: block;
  animation: fade-in-up 240ms ease;
}

.tax-species-list {
  display: flex;
  flex-wrap: wrap;
  gap: var(--space-xs);
  padding: var(--space-md) 0 var(--space-md) var(--space-lg);
  margin-left: var(--space-md);
  border-left: 1px dashed var(--border-default);
}

.tax-species-chip {
  display: inline-flex;
  align-items: center;
  padding: 4px 12px;
  background: rgba(255,255,255,0.03);
  border: 1px solid var(--border-subtle);
  border-radius: var(--radius-pill);
  font-family: var(--font-serif);
  font-style: italic;
  font-size: 0.88em;
  color: var(--ink-secondary);
  transition: all var(--t-fast);
  cursor: default;
}

.tax-species-chip.in-scope {
  cursor: pointer;
  border-color: var(--gold-soft);
  background: var(--gold-soft);
  color: var(--gold-bright);
}
.tax-species-chip.in-scope:hover {
  background: var(--gold);
  color: var(--bg-base);
  transform: translateY(-1px);
  box-shadow: var(--shadow-sm);
}

/* Compact version (scope) */
.tax-tree-compact .tax-node-header {
  grid-template-columns: 24px minmax(120px, 1fr) auto 60px;
}

/* === GALERÍA === */

.gallery {
  display: grid;
  grid-template-columns: repeat(auto-fill, minmax(220px, 1fr));
  gap: var(--space-md);
  width: 100%;
  max-width: 1280px;
  margin: var(--space-lg) auto 0;
}

.gallery-item {
  background: var(--bg-surface);
  border: 1px solid var(--border-subtle);
  border-radius: var(--radius-md);
  overflow: hidden;
  cursor: pointer;
  transition: all var(--t-normal);
  aspect-ratio: 4 / 5;
  position: relative;
}

.gallery-item:hover {
  transform: translateY(-3px);
  border-color: var(--border-accent);
  box-shadow: var(--shadow-lg);
}

.gallery-item img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  background: var(--bg-elevated);
}

.gallery-item-caption {
  position: absolute;
  inset: auto 0 0 0;
  padding: var(--space-md);
  background: linear-gradient(180deg, transparent 0%, rgba(0,0,0,0.85) 100%);
  color: var(--ink-primary);
  display: flex;
  flex-direction: column;
  gap: 2px;
}

.gallery-item-caption .name {
  font-family: var(--font-serif);
  font-style: italic;
  font-size: 1.1rem;
  font-weight: 600;
  color: var(--ink-primary);
  line-height: 1.1;
}
.gallery-item-caption .origin {
  font-size: 0.72rem;
  color: var(--gold-bright);
  letter-spacing: var(--tracking-wide);
  text-transform: uppercase;
  font-weight: 600;
}

.gallery-item.is-venezuelan::before {
  content: "VE";
  position: absolute;
  top: 10px;
  right: 10px;
  z-index: 2;
  background: var(--neon);
  color: var(--bg-base);
  padding: 3px 8px;
  border-radius: var(--radius-pill);
  font-size: 10px;
  font-weight: 700;
  letter-spacing: 0.1em;
}

/* === FICHA DE ESPECIE === */

.species-card {
  display: grid;
  grid-template-columns: 1.1fr 1fr;
  gap: var(--space-xl);
  align-items: stretch;
  height: 100%;
  padding: var(--space-lg) var(--space-2xl);
  max-width: 1380px;
  margin: 0 auto;
  box-sizing: border-box;
}

.species-image {
  border-radius: var(--radius-md);
  overflow: hidden;
  background: #000;
  display: grid;
  place-items: center;
  border: 1px solid var(--border-default);
  position: relative;
  width: 100%;
  height: 100%;
  min-height: 400px;
}
.species-image img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  background: #000;
}

.species-image .species-badge {
  position: absolute;
  top: var(--space-md);
  left: var(--space-md);
  padding: 4px 12px;
  background: var(--neon);
  color: var(--bg-base);
  border-radius: var(--radius-pill);
  font-size: 11px;
  font-weight: 700;
  letter-spacing: var(--tracking-widest);
}

.species-content {
  display: flex;
  flex-direction: column;
  gap: var(--space-md);
}

.species-eyebrow {
  font-family: var(--font-body);
  font-size: 0.75rem;
  font-weight: 600;
  letter-spacing: var(--tracking-widest);
  text-transform: uppercase;
  color: var(--gold);
}

.species-name {
  font-family: var(--font-serif);
  font-style: italic;
  font-weight: 500;
  font-size: clamp(2.2rem, 4.5vw, 3.6rem);
  line-height: 1;
  margin-bottom: 0;
}
.species-name .genus { color: var(--ink-tertiary); }
.species-name .epithet { color: var(--neon-bright); font-weight: 600; }

.species-tagline {
  color: var(--ink-tertiary);
  font-size: 0.9rem;
}

.species-fields {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: var(--space-md);
  margin-top: var(--space-sm);
}

.species-field {
  display: flex;
  flex-direction: column;
  gap: 4px;
}

.species-field .label {
  font-size: 0.7rem;
  font-weight: 600;
  letter-spacing: var(--tracking-wide);
  color: var(--gold);
  text-transform: uppercase;
}
.species-field .value {
  font-size: 0.92rem;
  color: var(--ink-secondary);
  line-height: var(--lh-snug);
}

.species-paragraphs {
  display: flex;
  flex-direction: column;
  gap: var(--space-sm);
  margin-top: var(--space-sm);
}

.species-paragraphs .label {
  font-size: 0.7rem;
  font-weight: 600;
  letter-spacing: var(--tracking-wide);
  color: var(--gold);
  text-transform: uppercase;
  margin-bottom: 4px;
}
.species-paragraphs p {
  font-size: 0.95rem;
  color: var(--ink-secondary);
  line-height: var(--lh-relaxed);
  margin: 0;
}

.species-thumbnails {
  margin-top: var(--space-md);
  display: flex;
  gap: var(--space-xs);
  flex-wrap: wrap;
}
.species-thumbnails .thumb {
  width: 56px;
  height: 56px;
  border-radius: var(--radius-sm);
  border: 1px solid var(--border-default);
  cursor: pointer;
  object-fit: cover;
  transition: all var(--t-fast);
}
.species-thumbnails .thumb:hover {
  transform: scale(1.08);
  border-color: var(--gold);
}

/* === LIGHTBOX === */

.lightbox-backdrop {
  position: fixed;
  inset: 0;
  background: rgba(5, 5, 8, 0.92);
  backdrop-filter: blur(8px);
  z-index: var(--z-modal);
  display: none;
  align-items: center;
  justify-content: center;
  padding: var(--space-xl);
  animation: fade-in-up 240ms;
}

.lightbox-backdrop.open {
  display: flex;
}

.lightbox-content {
  max-width: 92vw;
  max-height: 88vh;
  position: relative;
  display: flex;
  flex-direction: column;
  gap: var(--space-md);
  align-items: center;
}

.lightbox-content img {
  max-width: 100%;
  max-height: 78vh;
  object-fit: contain;
  border-radius: var(--radius-md);
  box-shadow: var(--shadow-lg);
}

.lightbox-close {
  position: fixed;
  top: 20px;
  right: 24px;
  background: var(--bg-surface);
  color: var(--ink-primary);
  border: 1px solid var(--border-default);
  width: 40px;
  height: 40px;
  border-radius: 50%;
  cursor: pointer;
  font-size: 22px;
  display: grid;
  place-items: center;
  transition: all var(--t-fast);
  font-family: inherit;
}
.lightbox-close:hover {
  background: var(--gold);
  color: var(--bg-base);
  border-color: var(--gold);
}

.lightbox-caption {
  color: var(--ink-secondary);
  font-family: var(--font-serif);
  font-style: italic;
  text-align: center;
  font-size: 1rem;
}

/* === COMPARATIVO === */

.compare {
  display: grid;
  grid-template-columns: 180px repeat(var(--cols, 4), 1fr);
  gap: 0;
  width: 100%;
  max-width: 1280px;
  margin: var(--space-md) auto;
  border-radius: var(--radius-md);
  overflow: hidden;
  border: 1px solid var(--border-default);
}

.compare-cell {
  padding: var(--space-md);
  background: var(--bg-surface);
  border-bottom: 1px solid var(--border-subtle);
  font-size: 0.85rem;
  line-height: var(--lh-snug);
  color: var(--ink-secondary);
}

.compare-cell.header {
  background: var(--bg-elevated);
  font-weight: 600;
  color: var(--ink-primary);
}

.compare-cell.species-header {
  background: var(--bg-base);
  text-align: center;
  font-family: var(--font-serif);
  font-style: italic;
  font-size: 1.2rem;
  font-weight: 600;
  color: var(--neon-bright);
  border-bottom: 2px solid var(--neon-soft);
  padding: var(--space-md);
}

.compare-cell.criterion {
  background: var(--bg-elevated);
  font-weight: 600;
  color: var(--gold);
  font-size: 0.78rem;
  text-transform: uppercase;
  letter-spacing: var(--tracking-wide);
}

/* === HÍBRIDOS === */

.hybrids-grid {
  display: grid;
  grid-template-columns: repeat(auto-fill, minmax(280px, 1fr));
  gap: var(--space-md);
  margin-top: var(--space-md);
}

.hybrid-card {
  background: var(--bg-surface);
  border: 1px solid var(--border-subtle);
  border-radius: var(--radius-md);
  padding: var(--space-md);
  border-left: 3px solid var(--gold);
}

.hybrid-card .name {
  font-family: var(--font-serif);
  font-style: italic;
  color: var(--ink-primary);
  font-weight: 600;
  margin-bottom: 4px;
}

.hybrid-card .parents {
  font-size: 0.85rem;
  color: var(--ink-tertiary);
  font-family: var(--font-serif);
  font-style: italic;
}

/* === CONTRAPORTADA === */

.outro {
  display: grid;
  place-items: center;
  min-height: 100%;
  height: 100%;
  width: 100%;
  text-align: center;
  padding: var(--space-xl);
  position: relative;
  box-sizing: border-box;
}

.outro-content {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: var(--space-lg);
  max-width: 720px;
}

.outro-thanks {
  font-family: var(--font-display);
  font-weight: 700;
  font-size: clamp(4rem, 9vw, 7rem);
  letter-spacing: -0.03em;
  color: var(--gold);
  text-shadow: 0 0 64px rgba(201,160,78,0.3);
}

.outro-credits {
  font-family: var(--font-serif);
  font-style: italic;
  color: var(--ink-secondary);
  font-size: 1.2rem;
  line-height: var(--lh-relaxed);
}

.outro-author {
  font-size: 1.4rem;
  font-weight: 600;
  color: var(--ink-primary);
}

.outro-logos {
  display: flex;
  gap: var(--space-2xl);
  align-items: center;
  margin-top: var(--space-md);
}
.outro-logos img {
  height: 48px;
  background: rgba(245,242,237,0.92);
  padding: 6px 10px;
  border-radius: var(--radius-sm);
}

/* === RESPONSIVE === */

@media (max-width: 900px) {
  .species-card {
    grid-template-columns: 1fr;
    grid-template-rows: 220px auto;
    gap: var(--space-md);
    padding: var(--space-md);
  }
  .species-fields { grid-template-columns: 1fr; }
  .compare {
    grid-template-columns: 100%;
    overflow-x: auto;
  }
  .gallery { grid-template-columns: repeat(auto-fill, minmax(150px, 1fr)); }
  .tax-node-header {
    grid-template-columns: 20px 1fr auto auto;
    font-size: 0.85rem;
  }
  .tax-node-meta { display: none; }
  .slide-pad { padding: var(--space-md); }
}

/* === SLIDE BACKGROUND PARA SECCIÓN === */

[data-section-color] {
  --section-color: var(--gold);
}
[data-section-color="cattleya"]     { --section-color: var(--sect-cattleya); }
[data-section-color="lawrenceanae"] { --section-color: var(--sect-lawrenceanae); }
[data-section-color="cattleyella"]  { --section-color: var(--sect-cattleyella); }
[data-section-color="maximae"]      { --section-color: var(--sect-maximae); }

/* Slim cinta superior por sección */
.slide-cinta {
  position: absolute;
  inset: 0 0 auto 0;
  height: 4px;
  background: var(--section-color, var(--gold));
  z-index: 5;
}

/* ========================================================================
   COMPLEMENTOS V1 (slides genéricos · intro · timeline · brand corner)
   ======================================================================== */

/* === HEADERS GENÉRICOS PARA SLIDES === */

.slide-eyebrow {
  display: inline-block;
  font-family: var(--font-body);
  font-size: 0.7rem;
  font-weight: 600;
  letter-spacing: var(--tracking-widest);
  text-transform: uppercase;
  color: var(--gold);
  margin-bottom: var(--space-sm);
}

.slide-title {
  font-family: var(--font-display);
  font-weight: 700;
  font-size: clamp(2rem, 4.5vw, 3.2rem);
  line-height: var(--lh-tight);
  letter-spacing: -0.02em;
  color: var(--ink-primary);
  margin: 0 0 var(--space-md) 0;
}

.slide-lead {
  font-family: var(--font-serif);
  font-style: italic;
  font-size: clamp(1rem, 1.6vw, 1.2rem);
  line-height: var(--lh-relaxed);
  color: var(--ink-secondary);
  max-width: 880px;
  margin: 0 0 var(--space-lg) 0;
}

/* === ITÁLICA CIENTÍFICA (alias) === */

.scientific {
  font-family: var(--font-serif);
  font-style: italic;
  font-weight: 500;
  letter-spacing: 0.005em;
}

/* === SLIDE INTRO (texto + asides) === */

.intro-grid {
  display: grid;
  grid-template-columns: 1.5fr 1fr;
  gap: var(--space-2xl);
  align-items: start;
  margin-top: var(--space-md);
}

.intro-text p {
  font-size: 1.05rem;
  line-height: var(--lh-relaxed);
  color: var(--ink-secondary);
  margin: 0 0 var(--space-md) 0;
}

.intro-aside {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: var(--space-md);
}

.aside-card {
  background: var(--bg-surface);
  border: 1px solid var(--border-subtle);
  border-left: 3px solid var(--gold);
  border-radius: var(--radius-md);
  padding: var(--space-md) var(--space-lg);
  display: flex;
  flex-direction: column;
  gap: 4px;
}

.aside-card-ven {
  border-left-color: var(--neon);
  background: linear-gradient(135deg, var(--bg-surface), rgba(196, 110, 224, 0.06));
}

.aside-num {
  font-family: var(--font-display);
  font-weight: 800;
  font-size: 2.4rem;
  line-height: 1;
  color: var(--ink-primary);
  letter-spacing: -0.03em;
}

.aside-card-ven .aside-num { color: var(--neon-bright); }

.aside-label {
  font-size: 0.72rem;
  font-weight: 500;
  letter-spacing: var(--tracking-wide);
  text-transform: uppercase;
  color: var(--ink-tertiary);
}

/* === FICHA DE ESPECIE — soporte para wrap, badge venezolana === */

.species-card.species-venezolana .species-name .epithet {
  color: var(--neon-bright);
  text-shadow: 0 0 32px rgba(196, 110, 224, 0.25);
}

.species-card > .species-image {
  border-left: 3px solid var(--accent-color, var(--gold));
}

.species-image[data-open-lightbox] { cursor: zoom-in; }
.species-image[data-open-lightbox]:hover img {
  transform: scale(1.02);
}
.species-image img {
  transition: transform var(--t-normal);
}

/* === COMPARE — cell wrapping === */

.compare-cell em {
  font-family: var(--font-serif);
  font-style: italic;
  color: var(--ink-primary);
}
.compare-cell.species-header em {
  color: var(--neon-bright);
}

/* === HÍBRIDOS — page reference === */

.hybrid-card .page-ref {
  font-size: 0.7rem;
  color: var(--ink-muted);
  letter-spacing: var(--tracking-wide);
  text-transform: uppercase;
  margin-top: var(--space-xs);
  font-family: var(--font-body);
  font-style: normal;
}

.hybrid-card .cross {
  color: var(--gold);
  font-style: normal;
  font-weight: 600;
  margin: 0 4px;
}

.hybrid-card .lineage {
  font-size: 0.8rem;
  color: var(--ink-tertiary);
  line-height: var(--lh-snug);
  display: block;
}

.hybrid-card .muted {
  color: var(--ink-muted);
  font-style: italic;
}

/* === HISTORY TIMELINE (quadricolor) === */

.history-timeline {
  list-style: none;
  padding: 0;
  margin: var(--space-md) 0 0 0;
  position: relative;
  display: flex;
  flex-direction: column;
  gap: var(--space-md);
}

.history-timeline::before {
  content: "";
  position: absolute;
  top: 8px;
  bottom: 8px;
  left: 100px;
  width: 1px;
  background: linear-gradient(180deg,
    transparent,
    var(--gold-soft) 10%,
    var(--gold-soft) 90%,
    transparent);
}

.hist-item {
  display: grid;
  grid-template-columns: 90px 1fr;
  gap: var(--space-xl);
  padding: var(--space-sm) 0;
  position: relative;
}

.hist-item::before {
  content: "";
  position: absolute;
  top: 22px;
  left: 96px;
  width: 9px;
  height: 9px;
  border-radius: 50%;
  background: var(--gold);
  box-shadow: 0 0 0 4px var(--bg-base), 0 0 16px var(--gold-soft);
}

.hist-year {
  font-family: var(--font-display);
  font-weight: 700;
  font-size: 1.4rem;
  color: var(--gold-bright);
  text-align: right;
}

.hist-content {
  display: flex;
  flex-direction: column;
  gap: 4px;
  padding-left: var(--space-md);
}

.hist-author {
  font-size: 0.78rem;
  font-weight: 600;
  letter-spacing: var(--tracking-wide);
  color: var(--ink-tertiary);
  text-transform: uppercase;
}

.hist-name {
  font-family: var(--font-serif);
  font-style: italic;
  font-size: 1.15rem;
  color: var(--ink-primary);
  font-weight: 500;
}

.hist-note {
  font-size: 0.85rem;
  color: var(--ink-secondary);
  line-height: var(--lh-snug);
  font-style: italic;
  font-family: var(--font-serif);
  margin-top: 4px;
}

/* === BRAND CORNER (firma fija arriba-derecha) === */

.brand-corner {
  position: fixed;
  top: 18px;
  right: 24px;
  z-index: 1000;
  display: flex;
  align-items: center;
  gap: var(--space-xs);
  font-family: var(--font-display);
  font-size: 0.72rem;
  font-weight: 500;
  letter-spacing: var(--tracking-wide);
  color: var(--ink-muted);
  pointer-events: none;
  opacity: 0.55;
  text-transform: uppercase;
}

.brand-corner-roman {
  font-family: var(--font-serif);
  font-style: italic;
  font-size: 1rem;
  color: var(--neon);
  font-weight: 500;
}

/* === RESPONSIVE === */

@media (max-width: 900px) {
  .intro-grid { grid-template-columns: 1fr; gap: var(--space-lg); }
  .intro-aside { grid-template-columns: 1fr 1fr; }

  .history-timeline::before { left: 70px; }
  .hist-item { grid-template-columns: 60px 1fr; gap: var(--space-md); }
  .hist-item::before { left: 65px; top: 18px; }
  .hist-year { font-size: 1.1rem; }

  .brand-corner { top: 10px; right: 12px; font-size: 0.6rem; }
  .brand-corner span:first-child { display: none; }
}

@media (max-width: 600px) {
  .intro-aside { grid-template-columns: 1fr; }
  .aside-num { font-size: 1.8rem; }
}